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sect. Trigynobrathys (Y.Kimura) N. Robson
Shrubs, subshrubs or perennial to annual herbs up to 2 m tall, the shrubs evergreen (or deciduous?), glabrous or very rarely with simple hairs, without dark glands, with taproot; branching pseudo-dichotomous or dichasial/monochasial or mixed, from up to 10 nodes. Stems 4-(rarely 6-) lined and compressed (ancipitous)when young and in distal nodes of herbaceous species, sooner or later becoming terete in shrubby species, densely glandular (when young in shrubby species); cortex exfoliating in strips or persistent; bark smooth. Leaves opposite or rarely 3-whorled, decussate, sessile, free to completely perfoliate, not forming interfoliar ridge or with base sheathing, deciduous at base or persistent (in herbaceous species); lamina entire, with venation parallel to pinnate or 1-nerved or secondarily with up to 7 basal veins, open? or closed, with tertiary venation laxly to densely reticulate or absent; laminar glands punctiform or rarely united in lines or striae; marginal gland dots dense, very small; ventral resin glands absent. Inflorescence 1-flowered with branching pseudo-dichotomous (usually from uppermost node) or 2-∞-flowered with branching dichasial/monochasial from uppermost node, often with subsidiary branches from lower nodes; bracts and bracteoles foliar or reduced. Flowers stellate, homostylous. Sepals 5(4), free, persistent, erect in fruit, with margin entire; veins 1-11; laminar glands wholly linear to mostly punctiform; marginal, submarginal, and inframarginal glands absent. Petals 5(4), persistent, with apiculus terminal or subterminal to obsolete, acute to acuminate or obtuse; margin entire; marginal glands absent; laminar glands wholly linear or distally interrupted to punctiform or absent. Stamen fascicles basically 5(4?), with 1-c. 80 stamens per fascicle, fascicles free or usually all united to form continuous or interrupted narrow ring of stamens or more rarely united 2 + 2 + 1 or stamens irregularly grouped, persistent; filaments very shortly united or apparently free; anthers yellow, gland amber; pollen type VIII. Ovary with (2)3-5(6-8) parietal placentae, ∞-ovulate; styles (2)3-5(6-8), free from bases contiguous; stigmas clavate to broadly capitate. Capsule (2)3-5-valved, coriaceous to chartaceous, with valves obscurely narrowly longitudinally vittate, sometimes with 1-3 flattish vesicles. Seeds narrowly cylindric, not carinate, without apical expansion; testa linear- scalariform to ribbed-scalariform or more rarely linear-reticulate to irregularly reticulate.