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sect. Roscyna (Spach) R. Keller
Perennial herbs up to 2 m tall, with stems erect or sometimes basally ascending from taproot, glabrous, without dark glands; branching lateral, confined to inflorescence or from up to all nodes. Stems 4-2-lined when young, becoming narrowly 4-winged to terete, eglandular. Leaves opposite, decussate, sessile to subsessile, free, persistent; lamina entire with venation pinnate, closed sometimes except lower 1-2 pairs, with tertiary reticulation dense, sometimes obscure; laminar glands pale, punctiform or shortly striiform, unequal; marginal gland dots dense, very small; ventral resin glands absent Inflorescence 1-c. 35-flowered with branching dichasial/monochasial from 1-7 nodes, often with subsidiary branches from lower nodes; bracts and bracteoles foliar or reduced, deciduous or persistent. Flowers stellate, homostylous. Sepals 5, free or united at base, persistent, spreading to deflexed in fruit, with margin entire; veins 5-17; laminar glands linear and distally interrupted to striiform or wholly punctiform; intramarginal glands few or absent. Petals 5, persistent and becoming reflexed, with apiculus present, lateral, or absent; margin entire; marginal glands absent; laminar glands linear, sometimes interrupted distally, numerous, or rarely absent. Stamen fascicles 5, free (or 4?, united 2+1+1+1), persistent, each single fascicle with c. 15-30 stamens; filaments united at base only; anthers yellow, gland amber; pollen type III. Ovary with central lacuna and 5(4) axile placentae, ¥-ovulate; styles 5(4), partly united or partly coherent or free; stigma narrowly to broadly capitate or infundibuliform. Capsule 5(4)-valved, coriaceous, with valves narrowly longitudinally vittate. Seeds cylindric, narrowly carinate to narrowly winged, sometimes with slight apical expansion; testa densely and shallowly linear-reticulate.