Hypericum caprifolium (Nomenclature)
Perennial herb 0.2-1 m tall, erect or ascending from creeping and rooting base, with woody taproot, basal branches few, diffuse, upper branches few, curved-ascending, or usually absent, rarely branched all down stem, the whole plant except inflorescence densely crisped-pubescent or (some leaves) puberulous. Stems reddish, wholly terete or sometimes (2)4(6)-lined in upper parts of inflorescence; internodes mostly exceeding leaves. Leaves sessile, all pairs except lowermost ones connate; lamina (15-)20-50 x (9-)12-24 mm, elliptic to oblong or ovate-oblong, subconcolorous, not glaucous, plane, spreading; apex rounded to obtuse, margin plane, base with sinus between pairs acute to obtuse or (in lowermost) cuneate; venation: 3 pairs of lateral curved-ascending from lower 0.25-0.5 of midrib; tertiary reticulation dense, plane not bullate/impressed; laminar glands pale, fairly dense, scattered, small, not prominent and sometimes black, sparse, scattered; intramarginal glands black, rather dense, irregular. Inflorescence c. 10-80-flowered from 1-4 nodes, densely curved-corymbiform to rounded-pyramidal, becoming monochasial after first flower, sometimes with flowering branches from up to 4 nodes, the whole then laxly cylindric; pedicels 3-5 mm; bracts and bracteoles linear-triangular, apex aristate, black-glandular-fimbriate, at least bracteoles with gland-fringed auricles. Flowers c. 12-15(?-20) mm in diam.; buds cylindric, subacute. Sepals 5-7 x 0.7-1.7 mm, subequal, narrowly lanceolate, aristate, with margin glandular-ciliate and arista gland-tipped; veins 5, branched and reticulating towards margin and distally; laminar glands pale, punctiform and (mostly) black, shortly linear to punctiform, often dense; marginal glands black, on short to long cilia. Petals bright? yellow, tinged or veined red, 8-11 x 3-4 mm, c. 1.6 x sepals, oblong-lanceolate to oblanceolate, rounded, without apiculus; laminar glands pale, punctiform to shortly striiform, distal; marginal glands black, in dense row, distal. Stamens (25?-)35-45, clearly 3-fascicled, longest 7-9 mm, c. 0.9 x petals; anther gland black. Ovary 3-locular, c. 2 x 1 mm, narrowly ovoid; styles 3, (3.5-)4-5 mm, 1.5-2.5 x ovary, spreading-incurved or spreading. Capsule 3.5-4 x c. 2.5 mm, ovoid, 0.6-0.75 x sepals, enclosed by petals twisting together. Seeds yellow-brown, c. 0.6 mm long; testa very finely scalariform (almost smooth).
2n=18 (Reynaud, 1986), 16 (Love & Kjellqvist, 1974); n=9 (Reynaud, 1986).
Streamsides, flushes and shaded places; 200-2000 m.
Spain (south-east: Andalucia, Murcia, Valencia, Castilla Nueva, Aragon).
H. caprifolium is easily distinguished from the North African H. naudinianum by the aristate sepals. From the scanty available evidence, the chromosome number is smaller (2n=16, not 18).